The other thing I notice is CORROSION on the saddles.
Here's the deal...some players will corrode the strings off a guitar in one set because their body chemistry is such that their sweat eats metal. Other guitarists can play for months on a set of strings and they're perfectly fine--usually because their hands don't sweat very much at all.
What you have going on there is galvanic corrosion caused by dissimilar metals. The brass saddle preferentially corrodes in electrical contact with the steel or nickel alloy of the guitar string. That creates a rough spot on the saddle that abrades the string through tuning and bending, which microscopically pulls the string over the saddle.
Graphite saddles will eliminate that problem. If you don't want to switch saddles, try a different lube, such as 3-in-1 oil. Use a very small dot of it, applied with the point of a toothpick. Do this after cleaning the corrosion off the saddles and polishing them.