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The original 5E3 used a 12AY7 for the preamp and a 12AX7 as a driver and a phase inverter. You should try it that way to see what you get, especially using input 2 for cleans. Or is that too clean for you? I have experience with the Weber 5E3 Proluxe, which is the same idea as your amp. Using 5881s, input 2 was just plain beautiful.
You're using a 12AT7 in V2 now, so a 12AY7 might sound a little brighter and a little quieter than the 12AT7--hard to say. You might also want to try a 12DW7 in V2. It's half 12AX7 and half 12AU7.
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