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Yes, but the 5751 goes between the 12AX7 and the 12AT7. It's about 80 percent of the 12AX7 and it has the same high plate impedance as the 12AX7, so it won't alter your tone (other than the normal difference among tube manufacturers).
The 12AT7 and 12AU7 are driver tubes. They have lower impedance and high current capabilities. The lower impedance generally causes some loss of highs. Some people like them in a preamp because they're mellow; others don't because they kill high end.
The 12AY7 has the third-highest impedance after the 12AX7 and 5751. It's a proper preamp tube. It's what the 5E3 was originally designed around.
The 12AV7 is really a switching tube. It's designed to drive relays and such at high currents and is optimized for being fully on or fully off.
Two 12AU7s would give you a practice amp--pretty darn quiet. No breakup.
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